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The mischievous Minions hope that Gru will return to a life of crime after the new boss of the Anti-Villain League fires him. Instead, Gru decides to remain retired and travel to Freedonia to meet his long-lost twin brother for the first time. The reunited siblings soon find themselves in an uneasy alliance to take down the elusive Balthazar Bratt, a former 1980s child star who seeks revenge against the world.

Evolving from single-celled yellow organisms at the dawn of time, Minions live to serve, but find themselves working for a continual series of unsuccessful masters, from T. Rex to Napoleon. Without a master to grovel for, the Minions fall into a deep depression. But one minion, Kevin, has a plan; accompanied by his pals Stuart and Bob, Kevin sets forth to find a new evil boss for his brethren to follow. Their search leads them to Scarlet Overkill, the world's first-ever super-villainess.